Tardigrade Taxonomy
 

Below is the current taxonomy of the Phylum Tardigrada down to the genus level. The taxonomy follows the Species Checklist by Degma, Guidetti & Bertolani (2009+). The Checklist is updated several times a year by Peter Degma and thus comprises an up to date list of all described valid tardigrade species. If you want to use this taxonomy, please cite the Checklist as requested in its Introduction.

 

The site was last updated on the 15-01-2020.

 

Phylum  Tardigrada  (Spallanzani, 1777)
        1. Class  Heterotardigrada  Marcus, 1927
               
1. Order  Arthrotardigrada  Marcus, 1927
                               
1. Family  Archechiniscidae  Binda, 1978
                                               
1. Genus  Archechiniscus  Schulz, 1953
                               
2. Family  Batillipedidae  Ramazzotti, 1962
                                               
1. Genus  Batillipes  Schulz, 1953
                               
3. Family  Coronarctidae  Renaud-Mornant, 1974
                                               
1. Genus  Coronarctus  Renaud-Mornant, 1974
                                               
2. Genus  Trogloarctus  Villora-Moreno, 19
96
                               
4. Family  Halechiniscidae  Thulin, 1928
                                       
1. Subfamily  Dipodarctinae  Pollock, 1995
                                               
1. Genus  Dipodarctus  Pollock, 1995
                                       
2. Subfamily  Euclavarctinae  Renaud-Mornant, 1983
                                               
1. Genus  Clavarctus  Renaud-Mornant, 1983
                                               
2. Genus  Euclavarctus  Renaud-Mornant, 1975
                                               
3. Genus  Exoclavarctus  Renaud-Mornant, 1983
                                               
4. Genus  Mobjergarctus  Bussau, 1982
                                               
5. Genus  Parmursa  Renaud-Mornant, 1984
                                               
6. Genus  Proclavarctus  Renaud-Mornant, 1983
                                       
3. Subfamily  Florarctinae  Renaud-Mornant, 1983
                                               
1. Genus  Florarctus  Delamare Deboutteville & Renaud-Mornant, 1965
                                               
2. Genus  Ligiarctus  Renaud-Mornant, 1982
                                               
3. Genus  Wingstrandarctus  Kristensen, 1984
                                       
4. Subfamily  Halechiniscinae  (Thulin, 1928)
                                               
1. Genus  Chrysoarctus  Renaud-Mornant, 1984
                                               
2. Genus  Halechiniscus  Richters, 1908
                                       
5. Subfamily  Orzeliscinae  (Schulz, 1963)
                                               
1. Genus  Mutaparadoxipus  Gross, Miller & Hochberg, 1952
                                               
2. Genus  Orzeliscus  du Bois-Reymond Marcus, 1952
                                               
3. Genus  Opydorscus  Renaud-Mornant, 1989
                                               
4. Genus  Paradoxipus  Kristensen & Higgins, 1989
                                       
6. Subfamily  Styraconyxinae  Kristensen & Renaud-Mornant, 1983
                                               
1. Genus  Angursa  Pollock, 1979
                                               
2. Genus  Bathyechiniscus  Steiner, 1926
                                               
3. Genus  Lepoarctus  Kristensen & Renaud-Mornant, 1983
                                               
4. Genus  Pleocola  Cantacuzène, 1951
                                               
5. Genus  Raiarctus  Renaud-Mornant, 1981
                                               
6. Genus  Rhomboarctus  Renaud-Mornant, 1984
                                               
7. Genus  Styraconyx  Thulin, 1942
                                               
8. Genus  Tetrakentron  Cuénot, 1892
                                               
9. Genus  Tholoarctus  Kristensen & Renaud-Mornant, 1983
                                       
7. Subfamily  Tanarctinae  Renaud-Mornant, 1980
                                               
1. Genus  Actinarctus  Schulz, 1935
                                               
2. Genus  Tanarctus  Renaud-Debyser, 1959
                                               
3. Genus  Zioella  Renaud-Mornant, 1987
                               
5. Family  Neoarctidae 
de Zio Grimaldi, D'Addabbo Gallo & Morone De Lucia, 1992
                                               
1. Genus  Neoarctus 
de Zio Grimaldi, D'Addabbo Gallo & Morone De Lucia, 1992
                               
6. Family  Neostygarctidae 
de Zio Grimaldi, D'Addabbo Gallo & Morone De Lucia, 1987
                                               
1. Genus  Neostygarctus 
de Zio Grimaldi, D'Addabbo Gallo & Morone De Lucia, 1982
                               
7. Family  Renaudarctidae 
Kristensen & Higgins, 1984
                                               
1. Genus  Renaudarctus 
Kristensen & Higgins, 1984
                               
8. Family  Stygarctidae 
Schulz, 1951
                                       
1. Subfamily  Megastygarctidinae 
Bello & de Zio Grimaldi, 1998
                                               
1. Genus  Megastygarcides 
McKirdy, Schmidt & McGinty-Bayly, 1976
                                       
2. Subfamily  Stygarctidinae 
Schulz, 1951
                                               
1. Genus  Faroestygarctus  Hansen, Kristensen & Jørgensen, 2012
                                               
2. Genus  Hansenarctus  Fujimoto & Ohtsuka, 2019
                                               
3. Genus  Mesostygarctus  Renaud-Mornant, 1979
                                               
4. Genus  Parastygarctus  Renaud-Debyser, 1965
                                               
5. Genus  Prostygarctus  Rubal, Veiga, Fontoura & Sousa-Pinto, 2013
                                               
6. Genus  Pseudostygarctus 
McKirdy, Schmidt & McGinty-Bayly, 1976
                                               
7. Genus  Stygarctus 
Schulz, 1951
               
2. Order  Echiniscoidea  Richters, 1926
                               
1. Family  Echiniscoididae  Krsitensen & Hallas, 1980
                                       
1. Subfamily  Echiniscoidinae  Møbjerg, Kristensen & Jørgensen, 2016
                                               
1. Genus  Anisonyches  Pollock, 1975
                                               
2. Genus  Echiniscoides  Plate, 1888
                                       
2. Subfamily  Isoechiniscoidinae  Møbjerg, Kristensen & Jørgensen, 2016
                                               
1. Genus  Isoechiniscoides  Møbjerg, Kristensen & Jørgensen, 2016
                               
2. Family  Carphaniidae  Binda & Krsitensen, 1986
                                               
1. Genus  Carphania  Binda, 1978
                               
3. Family  Oreellidae  Puglia, 1959
                                               
1. Genus  Oreella  Murray, 1910
                               
4. Family  Echiniscidae  Thulin, 1928
                                               
1. Genus  Acanthechiniscus  Vecchi, Cesari, Bertolani, Jönsson, Rebecchi & Guidetti, 2016
                                               
2. Genus  Antechiniscus  Kristensen, 1987
                                               
3. Genus  Barbaria  Michalczyk, Gısiorek, Morek & Stec, 2019
                                               
4. Genus  Bryochoerus  Marcus, 1936
                                               
5. Genus  Bryodelphax  Thulin, 1928
                                               
6. Genus  Claxtonia  Gısiorek & Michalczyk, 2019
                                               
7. Genus  Cornechiniscus  Maucci & Ramazzotti, 1981
                                               
8. Genus  Diploechiniscus  Vicente, Fontoura, Cesari, Rebecchi, Guidetti, Serrano & Bertolani, 2013
                                               
9. Genus  Echiniscus  Schultze, 1840
                                              
10. Genus  Hypechiniscus  Thulin, 1928
                                              
11. Genus  Kristenseniscus  Gısiorek, Morek, Stec & Michalczyk, 2019
                                              
12. Genus  Mopsechiniscus  du Bois-Reymond Marcus, 1944
                                              
13. Genus  Multipseudechiniscus  Schulte & Miller, 2011
                                              
14. Genus  Nebularmis  Gısiorek & Michalczyk, 2019
                                              
15. Genus  Novechiniscus  Kristensen, 1987
                                              
16. Genus  Parechiniscus  Cuénot, 1926
                                             
17. Genus  Proechiniscus  Kristensen, 1987
                                              
18. Genus  Pseudechiniscus  Thulin, 1911
                                              
19. Genus  Stellariscus  Gısiorek, Suzuki, Kristensen & Michalczyk, 2018
                                              
20. Genus  Testechiniscus  Kristensen, 1987
                                              
21. Genus  Viridiscus  Gısiorek & Michalczyk, 2019
        2. Class  Mesotardigrada  Rahm, 1937
               
1. Order  Thermozodia  Ramazzotti & Maucci, 1983
                               
1. Family  Thermozodiidae  Rahm, 1937
                                               
1. Genus  Thermozodium  Rahm, 1937
        3. Class  Eutardigrada  Richters, 1926
               
1. Order  Apochela  Schuster, Nelson, Grigarick & Christenberry, 1980
                               
1. Family  Milnesiidae  Ramazzotti, 1962
                                               
1. Genus  Bergtrollus  Dastych, 2011
                                               
2. Genus  Limmenius  Horning, Schuster & Grigarick, 1978
                                               
3. Genus  Milnesioides  Claxton, 1999
                                               
4. Genus  Milnesium  Doyère, 1840
               
2. Order  Parachela  Schuster, Nelson, Grigarick & Christenberry, 1980
                       
1. Superfamily  Eohypsibioidea  Bertolani & Kristensen, 1987
                               
1. Family  Eohypsibiidae  Bertolani & Kristensen, 1987
                                               
1. Genus  Austeruseus 
Trygvadóttir & Kristensen, 2011
                                               
2. Genus  Bertolanius 
Özdikmen, 2008
                                               
3. Genus  Eohypsibius 
Kristensen, 1982
                        2. Superfamily  Hypsibioidea  Pilato, 1969
                               
1. Family  Calohypsibiidae  Pilato, 1969
                                               
1. Genus  Calohypsibius 
Thulin, 1928
                               
2. Family  Hypsibiidae  Pilato, 1969
                                       
1. Subfamily  Diphasconinae  Dastych, 1992
                                               
1. Genus  Diphascon 
Plate, 1889
                                        2. Subfamily  Hypsibiinae  Pilato, 1969
                                               
1. Genus  Borealibius 
Pilato, Guidetti, Rebecchi, Lisi, Hansen & Bertolani, 2006
                                                2. Genus  Hypsibius  Ehrenberg, 1848
                                        3. Subfamily  Itaquasconinae  Rudescu, 1964
                                                1. Genus  Adropion  Pilato, 1987
                                                2. Genus  Astatumen  Pilato, 1997
                                                3. Genus  Bindius  Pilato, 2009
                                                4. Genus  Itaquascon  de Barros, 1939
                                                5. Genus  Mesocrista  Pilato, 1987
                                                6. Genus  Parascon  Pilato & Binda, 1987
                                                7. Genus  Platicrista  Pilato, 1987
                                               
8. Genus  Sarascon  Guil, Rodrigo & Machordom, 2015
                                       
4. Subfamily  Pilatobiinae  Bertolani, Guidetti, Marchioro, Altiero, Rebecchi & Cesari, 2014
                                                1. Genus  Pilatobius  Bertolani, Guidetti, Marchioro, Altiero, Rebecchi & Cesari, 2014
                                        ?. Subfamily  ?
                                               
1. Genus  Acutuncus  Pilato & Binda, 1997

                                                2. Genus  Mixibius  Pilato, 1992
                                3. Family  Microhypsibiidae  Pilato, 1998
                                                1. Genus  Microhypsibius  Thulin, 1928
                                4. Family  Ramazzottiidae  Sands, McInnes, Marley, Goodall-Copestake, Convey & Linse, 2008
                                               
1. Genus  Cryoconicus 
Zawierucha, Stec, Lachowska-Cierlik, Takeuchi, Li & Michalczyk, 2018
                                                2. Genus  Hebesuncus  Pilato, 1987
                                                3. Genus  Ramazzottius  Binda & Pilato, 1986
                        3. Superfamily  Isohypsibioidea  Sands, McInnes, Marley, Goodall-Copestake, Convey & Linse, 2008
                               
1. Family  Doryphoribiidae  Gısiorek, Stec, Morek & Michalczyk, 2019
                                                1. Genus  Apodibius  Dastych, 1983
                                                2. Genus  Doryphoribius  Pilato, 1969
                                                3. Genus  Grevenius  Gısiorek, Stec, Morek & Michalczyk, 2019
                                                4. Genus  Paradiphascon  Dastych, 1992
                                               
5. Genus  Pseudobiotus 
Nelson, 1980
                                                6. Genus  Thulinius  Bertolani, 2003
                               
2. Family  Halobiotidae  Gısiorek, Stec, Morek & Michalczyk, 2019
                                               
1. Genus  Halobiotus 
Kristensen, 1982
                               
3. Family  Hexapodibiidae  Cesari, Vecchi, Palmer, Bertolani, Pilato, Rebecchi & Guidetti, 2016
                                               
1. Genus  Haplohexapodibius 
Pilato & Beasley, 1987
                                               
2. Genus  Haplomacrobiotus  May, 1948
                                               
3. Genus  Hexapodibius  Pilato, 1969
                                               
4. Genus  Parhexapodibius  Pilato, 1969
                                4. Family  Isohypsibiidae  Sands, McInnes, Marley, Goodall-Copestake, Convey & Linse, 2008
                                                1. Genus  Dastychius  Pilato, 2013
                                               
2. Genus  Dianea  Gısiorek, Stec, Morek & Michalczyk, 2019
                                                3. Genus  Eremobiotus  Biserov, 1992
                                                4. Genus  Fractonotus  Pilato, 1998
                                                5. Genus  Isohypsibius  Thulin, 1928
                                               
6. Genus  Ramajendas 
Pilato & Binda, 1990
                                                7. Genus  Thalerius  Dastych, 2009
                                                8. Genus  Ursulinius  Gısiorek, Stec, Morek & Michalczyk, 2019
                        4. Superfamily  Macrobiotoidea  Thulin, 1928
                               
1. Family  Macrobiotidae  Thulin, 1928
                                                1. Genus  Biserovus  Guidetti & Pilato, 2003
                                                2. Genus  Calcarobiotus  Dastych, 1993
                                                       
1. Subgenus  Calcarobiotus  Dastych, 1993
                                                       
2. Subgenus  Discrepunguis  Guidetti & Bertolani, 2001
                                                3. Genus  Famelobiotus  Pilato, Binda & Lisi, 2004
                                                4. Genus  Insuetifurca  Guidetti & Pilato, 2003
                                                5. Genus  Macrobiotus  Schultze, 1834
                                                6. Genus  Mesobiotus  Vecchi, Cesari, Bertolani, Jönsson, Rebecchi & Guidetti, 2016
                                                7. Genus  Minibiotus  Schuster, 1980
                                                8. Genus  Minilentus  Guidetti & Pilato, 2003
                                                9. Genus  Paramacrobiotus  Guidetti, Schill, Bertolani, Dandekar & Wolf, 2009
                                              10. Genus  Pseudodiphascon  Ramazzotti, 1965 (genus dubius)
                                              11. Genus  Pseudohexapodibius  Bertolani & Biserov, 1996
                                              12. Genus  Schusterius  Kaczmarek & Michalczyk, 2006
                                              13. Genus  Tenuibiotus  Pilato & Lisi, 2011
                                              14. Genus  Xerobiotus  Bertolani & Biserov, 1996
                                2. Family  Murrayidae  Guidetti, Gandolfi, Rossi & Bertolani, 2005
                                               
1. Genus  Dactylobiotus 
Schuster, 1980
                                                2. Genus  Macroversum  Pilato & Catanzaro, 1988
                                                3. Genus  Murrayon  Bertolani & Pilato, 1988
                                3. Family  Richtersiidae  Guidetti, Rebecchi, Bertolani, Jönsson, Kristensen & Cesari, 2016
                                               
1. Genus  Adorybiotus 
Maucci & Ramazzotti, 1981
                                                2. Genus  Diaforobiotus  Guidetti, Rebecchi, Bertolani, Jönsson, Kristensen & Cesari, 2016
                                                3. Genus  Richtersius  Pilato & Binda, 1989

                        ?. Superfamily  ?
                                ?. Family  Necopinatidae  Ramazzotti & Maucci, 1983
                                               
1. Genus  Necopinatum 
Pilato, 1971
 

 

 

 

 

 

 

 

 


Copyright by £ukasz Michalczyk (2012-2019)

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